Sac Fungi are part of the phylum Ascomycota, which includes the ascomycetes. With around 64,000 species, they are one of the largest phyla of the kingdom fungus.
Sac-fungi are what they are often referred to as. They are a part of the sub-kingdom of Dikarya (presence of dikaryon). Ascospores are non-motile spores produced by ascobacteria that are sexually reproducing. They are formed in a sac-like structure known as an ascus, which is a sac-like structure. Each ascus contains 4-8 ascospores, depending on the species.
A wide variety of habitats and eating habits are found in Ascomycetes; some are saprophytes, whilst others are pathogens that cause a variety of diseases in both animals and plants. Some ascomycetes, such as shiitake mushrooms, are edible. Some of them, such as lichens and mycorrhizae, coexist in a symbiotic relationship with one another.
Yeast, powdery mildews, cup fungi, morels, truffles, Neurospora, Aspergillus, Cladonia, Penicillium, Candida, Claviceps, and other ascomycetes are examples of ascomycetes that are commonly encountered.
Ascomycetes are extremely significant to the economy. It is from fermentation that we obtain fermented foods (bread, cheese, alcoholic beverages), antibiotics (Penicillin), and a variety of compounds. In biological study and research, a wide variety of species are used (yeast, Neurospora). Morels and truffles are delicacies that are enjoyed by many.
Ascomycetes are further subdivided based on the diverse structures of asci carrying structures and the different methods of releasing ascospores from these structures.
Characteristics of the Ascomycetes and Sac Fungi
- The ascus or asci, which is a reproductive structure seen in the majority of ascomycetes, is a distinguishing feature of the group.
- They are primarily terrestrial, parasitic, or coprophilous in nature.
- They can be either unicellular or multicellular fungus, depending on their size.
Four, the mycelium is made up of septate and branching hyphae (fungal filaments).
- Chitin or -glucans are the building blocks of the cell wall.
- Because of the presence of septal holes, there is cytoplasmic continuity.
- Asexual reproduction occurs through the generation of conidia on conidiophores that are produced exogenously.
- Yeast reproduces asexually through the process of budding.
- The process of sexual reproduction occurs by conjugation between two gametangias. They fall into one of two categories: homothallic or heterothallic.
- The ascocarp is the term used to describe the fruiting body. Ascocarps are classified into four types:
- a) Cleistothecium -When a fruiting body is spherical and securely closed, as in the case of Aspergillus, it is called a cleistothecium.
- b) Perithecium- The fruiting body is flask-shaped with one exterior opening, like in Neurospora, and has a single internal opening.
- c) Apothecium-For example, the fruiting body of an Apothecium plant is cup-shaped, and asci are present in the hymenium, e.g. Peziza.
- d) Ascostroma- There is no distinguishable fruiting body on this species. Mycosphaerella, for example, is an asci that can be found in the stroma.
- In ascii, ascospores are created by the organism itself.
Sexual reproduction of Ascomycetes :-
- Two different types of mating hyphae come together and fuse together.
The occurrence of plasmogamy occurs, but it does not immediately follow karyogamy.
- The fused structure has two haploid nuclei from each parent, which is referred to as dikaryon nuclei.
- With the help of dikaryotic cells, new hyphae are formed.
- Asci grows at the tip of the hyphae in the ascocarp.
- The nuclei of each ascus fuse together (karyogamy), resulting in the formation of an ascus diploid zygote.
- Formation of Ascospores: The diploid zygote undergoes meiosis, resulting in the formation of four haploid nuclei, which then undergo mitotic division, resulting in the formation of eight haploid nuclei. Each nucleus has a large amount of cytoplasm, which is surrounded by a strong cell wall. These are referred to as cospores.
- A pore, a slit, or a hinged lid in the asci allows ascospores to be freed from the asci and distributed by the wind.
- Ascospores germinate to create new mycelia in the presence of favourable circumstances.
The Importance of Ascomycetes in the Economy
The Ascomycota are the largest category of fungi, and they have a wide range of habitats. These people are cosmopolitan, and they provide us with a variety of beneficial items. They provide us with a variety of meals, medications, and chemical products.
- Yeast is essential for the fermentation of fermented products in the brewing, bread, and cheese industries.
Penicillin, the antibiotic obtained from Penicillium chrysogenum, is well-known to most people.
- A fungus called Tolypocladium niveum produces the immunosuppressant cyclosporin, which is used to treat cancer. It is employed in the treatment of organ transplantation and autoimmune illnesses.
- Ascomycetes are responsible for the production of a wide range of organic acids and enzymes, including citric acid, gluconic acid, amylases, proteases, and other enzymes.
- Fifth, the herb Claviceps purpurea (Ergot) is used as a medication to reduce excessive bleeding during menstrual periods and to speed up the process of childbirth.
- Various types of cheese are made from different Penicillium species, such as Camembert, Brie, Roquefort, and other similar varieties.
- The fungus Aspergillus is utilised in the production of soy sauce as well as the production of various Asian alcoholic beverages.
- Morels, truffles, and lobster mushrooms are all considered delicacies of the fungus world.
- Microorganisms such as Neurospora, Saccharomyces, and others are frequently used in genetics research.
Conclusion
There are three major classes of pathogenic Ascomycota at this time: the Saccharomycotina, the Taphrinomycotina, and the Pezizomycotina. These three primary classes account for all of the pathogenic members of Class Ascomycota.
Characteristics of the Ascomycetes and Sac Fungi
- The ascus or asci, a reproductive structure seen in the majority of ascomycetes, is a distinguishing characteristic of the group.
- They are primarily terrestrial, parasitic, or coprophilous in nature.
- They can be classified as either unicellular or multicellular fungus.
- The mycelium is made up of septate and branching hyphae, which are arranged in a spiral pattern.
Ascomycetes can be found in every form of habitat, including freshwater and marine habitats, tropical and temperate forests, and extreme conditions such as deserts. Ascomycetes can be found in every type of habitat. Many organisms play a significant part in the decomposition of organic matter.